Abdullah MT. 2003. Biogeography and variation of Cynopterus brachyotis in Southeast Asia. PhD thesis. The University of Queensland, St Lucia, Australia.
Corbet, GB, Hill JE. 1992. The mammals of the Indomalayan region: a systematic review. Oxford University Press, Oxford.
Hall LS, Gordon G. Grigg, Craig Moritz, Besar Ketol, Isa Sait, Wahab Marni and M.T. Abdullah. 2004. Biogeography of fruit bats in Southeast Asia. Sarawak Museum Journal LX(81):191-284.
Wilson DE, Reeder DM. 2005. Mammal species of the world. Smithsonian Institution Press, Washington DC.
